The Whitewings are an order of Pegasus Knights in service to the kingdom of Macedon. Led by the Macedonian princess, Minerva, its members are three sisters: Palla, Catria, and Est. Like Minerva, they join Marth’s army after Macedon falls to darkness during the War of Shadows. Later, Est was taken captive to the neighboring continent of Valentia, where Palla and Catria follow to search for their lost sister.